Due to my " upgrade existing system" I'm using ext4 and cannot see or feel any real difference over btrfs or reiser4 (i've got both systems dual booting on the same machine).
So essentially I'm not sure but do think (due to the articles @Thanatermesis posted) reiser4 is the better option.
About @yoda 's USB experience: I'm not so sure reiser4 itself is to blame, could be the disk not getting registered correctly as a high speed device. That might be caused by reiser4 being there and the firmware not knowing what to do with it.
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